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This was about 2 minutes in Lightroom using the masking brush with Auto Mask turned on and just increasing the exposure of the areas to turn them white. You could do a LOT better if you took more time and effort. The file is small and I think a large scan would work a lot better. I set the brush to MAXIMUM size and just had to do about 8 to 10 clicks to chose 4 areas.
You just have to be careful to select spots with all densities you want to eliminate.
I really need to learn Lightroom better. This is motivation. I had no idea you could do this!