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Your footage is probably sitting on a sub-pixel increment (assuming that your comp viewer settings are at full quality, and your layers are not set to draft). Do your positional values in your timeline have non-integer (decimal point) values?
Also check your comp resolution settings. If they are set to manual and do not inherit the parent comp settings (maybe you turned off that option in the prefs), the pre-comp will retain its (possibly reduced) preview resolution. Likewise, some layer switches may affect the result.
usually this phenomenon happened to me if i have used 3D compositions that i´ve precomposed and objects became blurry.
i then was in a hurry but i belive i solved it by deactivating the 3D switch my precomp and just activated "collapse transformation."
I did not changed anything in my master comp.
By just activating "collapse transformations" solved it for me. It's odd to me why though.
My money is on Mylenium's hypothesis. "Preserve resolution when nested" doesn't do what I suspect you think it does, Christopher. Ordinarily, when you switch a comp's preview resolution from, say, 50% to 100%, AE's renderer renders all nested comps at that same resolution. When you check "preserve resolution when nested", AE doesn't switch the nested comp's resolution, meaning any comps that are set to preview at a lower resolution won't be switched to 100% resolution if you decide to preview the top level comp at 100%.
Collapsing transformations might do something more like what you thought "Preserve Resolution" does. It forces the AE renderer to rasterize the contents of a nested comp inside the main comp. This means there is no intermediate rasterization of the nested comp, so the "Preserve resolution" setting is completely ignored.